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The park had been deemed a success story for elephant conservation, as well as being a refuge for endangered species such as tigers and rhinos.
Officials said a lack of resources allowed the fires to spread rapidly.
"We have not been able to assess the loss immediately because the fire is still raging at some places," the park's chief conservation officer Tika Ram Adhikari told BBC News.
"But given our past experiences, our estimate is that around 40% of small mammals, 60% of insects and significant number of birds have been lost in the fire."
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